Woods Hole
ATLAS - GAZETTEER COL

NOAA Technical Memorandum NMFS

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AUGUST 1987

ICHTHYOPLANKTON AND STATION DATA FOR

CALIFORNIA COOPERATIVE OCEANIC FISHERIES

INVESTIGATIONS SURVEY CRUISES IN 1951

David A. Ambrose, Richard L. Charter,
H. Geoffrey Moser and Celeste R. Santos Methot

PLEASE RETURN

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INSTITUTION DATA LIBRARY

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NOAA-TM-NMFS-SWFC- 79

U.S. DEPARTMENT OF COMMERCE

National Oceanic and Atmospheric Administration

National Marine Fisheries Service

Southwest Fisheries Center

NOAA Technical Memorandum NMFS

The National Oceanic and Atmospheric Administration (NOAA), organized in
1970, has evolved into an agency which establishes national policies and
manages and conserves our oceanic, coastal, and atmospheric resources. An
organizational element within NOAA, the Office of Fisheries is responsible for
fisheries policy and the direction of the National Marine Fisheries Service
(NMFS).

In addition to its formal publications, the NMFS uses the NOAA Technical
Memorandum series to issue informal scientific and technical publications when
complete formal review and editorial processing are not appropriate or feasible.
Documents within this series, however, reflect sound professional work and may
be referenced in the formal scientific and technical literature.

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Woods Hole Oceanographlc Institution
ATLAS • GAZETTEER COLLECTION

NOAA Technical Memorandum NMFS

This TM series is used lor docu menial ion and iimeiy communication ot preliminary results, imtenm reports, or
special purpose information, and have not received complete formal review, editorial control, or detailed editing

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AUGUST 1987

ICHTHYOPLANKTON AND STATION DATA FOR

CALIFORNIA COOPERATIVE OCEANIC FISHERIES

INVESTIGATIONS SURVEY CRUISES IN 1951

David A. Ambrose, Richard L. Charter,
H. Geoffrey Moser and Celeste R. Santos Methot
Southwest Fisheries Center
National Marine Fisheries Service, NOAA
La Jolla, California 92038

NOAA-TM-NMFS-SWFC-79

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U.S. DEPARTMENT OF COMMERCE

Malcolm Baldrige. Secretary

National Oceanic and Atmospheric Administration

Anthony J. Calio. Administrator

National Marine Fisheries Service

William E. Evans, Assistant Administrator for Fisheries

CONTENTS

Page

List of Figures iii

List of Tables iv

Abstract 1

Introduction 1

Sampling Area and Pattern 2

Sampling Gear and Methods 3

Laboratory Procedures 4

Identification 5

Computer Entry and Editing 10

Species Summary 10

Explanation of Tables 11

Acknowledgments 12

Literature Cited 13

Figures 15

Tables 29

Index 192

li

LIST OF FIGURES

Page

Figure 1. Composite arrangement of diagrammatic
charts showing areas sampled on each

CalCOFI cruise during 1951 15

Figure 2. Station pattern for CalCOFI Cruise 5101

showing tracks for each vessel 16

Figure 3. Station pattern for CalCOFI Cruise 5102 17

Figure 4. Station pattern for CalCOFI Cruise 5103 18

Figure 5. Station pattern for CalCOFI Cruise 5104 19

Figure 6. Station pattern for CalCOFI Cruise 5105 20

Figure 7. Station pattern for CalCOFI Cruise 5106 21

Figure 8. Station pattern for CalCOFI Cruise 5107 22

Figure 9. Station pattern for CalCOFI Cruise 5108 23

Figure 10. Station pattern for CalCOFI Cruise 5109 24

Figure 11. Station pattern for CalCOFI Cruise 5110 25

Figure 12. Station pattern for CalCOFI Cruise 5111 26

Figure 13. Station pattern for CalCOFI Cruise 5112 27

Figure 14. The basic station plan for CalCOFI

cruises from 1950 to the present 28

in

LIST OF TABLES

Page

Table 1. Station and plankton tow data for

CalCOFI cruises in 1951 29

Table 2. Pooled occurrences of fish larvae taken

during CalCOFI cruises in 1951 65

Table 3. Pooled numbers of fish larvae taken

during CalCOFI cruises in 1951 68

Table 4 . Numbers of fish larvae taken on stations

occupied during CalCOFI cruises in 1951 71

Table 5. Summary of pooled occurrences of fish
larvae taken on CalCOFI cruises from
1951-1960 187

Table 6. List of stations which were occupied

twice in one month during 1951 191

IV

ABSTRACT

This report provides ichthyoplankton and associated station
and tow data from California Cooperative Oceanic Fisheries
Investigations (CalCOFI) cruises conducted off California and
Baja California in 1951. It is the first report in a series that
presents these data for all biological-oceanographic CalCOFI
surveys from 1951 to the present. A total of 1368 stations were
occupied during 12 monthly multivessel cruises over the quarter-
million square mile survey area which extends from the
California-Oregon border to Cape San Lucas, Mexico and seaward to
several hundred miles. The data are listed in a series of 6
tables; the background, methodology, and information necessary
for interpretation and quantitative analysis of the data are
presented in an accompanying text. All pertinent station and tow
data, including volumes of water strained and standard haul
factors are listed in the first table. Another key table lists,
by station and month, standardized counts of each of the 134
larval fish taxa identified from survey samples. This and
subsequent reports will make the CalCOFI ichthyoplankton and
station data available to all investigators and will serve as
guides to the newly developed computer data base.

INTRODUCTION

This report, the first of a series, provides ichthyoplankton
and associated station and tow data from California Cooperative
Oceanic Fisheries Investigations (CalCOFI) joint biological-
oceanographic survey cruises conducted in 1951. This program
was initiated in 1949 under the sponsorship of the Marine
Research Committee of the State of California, to study the
population fluctuations of the Pacific sardine (Sardinops sagax)
and the environmental factors that may play a role in such
fluctuations. CalCOFI, known as the California Cooperative
Sardine Research Program from 1949 to 1953, was made up of
representatives of the South Pacific Fisheries Investigations
(SPFI) of the U.S. Fish and Wildlife Service [now the La Jolla
Laboratory, National Marine Fisheries Service (NMFS) ] , the
Scripps Institution of Oceanography (SIO) , the California
Department of Fish and Game (CDFG) , the California Academy of
Sciences (CAS) and the Hopkins Marine Station of Stanford
University. The first three of these agencies supplied ships and
personnel to conduct the sea surveys. NMFS processed the
plankton samples and analyzed the ichthyoplankton from them. SIO
processed and analyzed the hydrographic samples and measurements
and also analyzed invertebrate groups from the plankton samples.

The boundaries, station placement, and sampling frequency
for the CalCOFI survey area were based on the results of joint
biological and oceanographic cruises conducted by NMFS and SIO
during 1939-41. Those cruises were designed to collect sardine
eggs and larvae and associated hydrographic data over the entire
areal and seasonal spawning range of the species. On these
survey cruises, plankton tows were made to 70 m, a depth which

encompassed the vertical distribution of sardine eggs and larvae.
Wide-ranging joint biological and oceanographic survey cruises
were resumed in 1949 with sardine as the focus; however, an
increasing interest in other biological components resulted in
the deepening of standard tows to 140 m in 1951. This marked the
beginning of truly guantitative ichthyoplankton sampling on
CalCOFI surveys.

Data resulting from CalCOFI surveys in 1951 have been
published in a number of forms. Hydrographic data (Reid et al . ,
1963), zooplankton volumes (Staff, SPFI, 1952; Thrailkill, 1956;
Smith, 1971) and ichthyoplankton data for selected species
(Ahlstrom, 1953) were presented in standard formats. The latter
lists counts for eggs and larvae of sardine and for larvae of
northern anchovy (Engraulis mordax) , jack mackerel {Trachurus
symmetricus) , Pacific mackerel {Scomber japonicus) , Pacific hake
(Merluccius productus) , and rockfishes (Sebastes spp.). Also,
length freguencies are listed for sardine and anchovy larvae.
Distribution maps of larvae of 5 of these species taken on
CalCOFI surveys during 1951 are presented in the CalCOFI Atlas
series (Kramer and Ahlstrom, 1968; Ahlstrom, 1969; Kramer, 1970;
Ahlstrom et al., 1978).

A computer data base for eggs and larvae of sardine and
anchovy and for larvae of hake, and the two mackerels was
established in 1969. The development of a data base for other
fish larvae is a complex undertaking because competency of
identification has evolved steadily over the past 38 years. We
began the task of producing a CalCOFI ichthyoplankton data base
and associated data report series in 1983. All available
original records for 1951 were subjected to an extensive
verification and editing process to produce this report. This
and subseguent reports make the CalCOFI ichthyoplankton and
station data available to all investigators and serve as guides
to the computer data base. The data base will be modified when
additional errors are discovered and when composite taxa from the
earlier years are reidentif ied. These reports are the
fundamental reference documents against which subsequent changes
in the data base can be compared.

SAMPLING AREA AND PATTERN

In 1951, CalCOFI survey cruises were conducted at monthly
intervals. A total of 1368 stations were occupied on 12
cruises, with an average of 120 stations per cruise (range of 65-
170) . Coverage of the survey station pattern varied among
cruises and the entire quarter-million square mile survey area
was not covered on any single cruise (Figures 1-13; Table 1) .
The area off northern California (lines 40-57) was covered on
only two cruises made in July and August. Coverage off central
California (lines 60-77) was more consistent with stations
occupied monthly, except during February and March. The area
between Pt. Conception, California and Pt. San Juanico, Baja
California (lines 80-137) was surveyed monthly, except for

December when coverage extended south only to San Cristobal Bay
(line 123) . The area off southern Baja California (lines 140-
157) was surveyed in March, June, and September. Coverage
extended seaward to stations 120 or 130 (approximately 300-400
miles offshore) on some lines in the main sampling area but
typically did not extend beyond station 90 (approximately 160-250
miles offshore) . Offshore coverage was greatest between January
and July and generally diminished during later cruises.

Six vessels were employed on these cruises: the Black
Douglas, of NMFS, the Crest, Horizon, and Paolina T of SIO, and
the N. B. Sco field and Yellowfin of CDFG. Two to four vessels
participated on each cruise with three being the usual number.
The Crest was used on all cruises except 5112 and the Black
Douglas on all but 5101 and 5112. The other four vessels
participated on a total of 13 cruises (Ahlstrom, 1953) .

SAMPLING GEAR AND METHODS

The standard CalCOFI net used from 1949 to 1969 had a 1-m
diameter mouth opening (0.785 m 2 area) and an overall length of
about 5 m. The net was constructed of 3 0xxx gauze, a heavy duty
grade of silk bolting cloth, with a mesh size of 0.55 mm after
shrinkage. The last 40 cm of the cone and the cod end were
constructed of 56xxx grit gauze which had a mesh size of 0.25 mm
after shrinkage. The net ring was fastened to a short 3-lead
bridle connected to several meters of line which attached to the
towing cable by a clamp. A current meter was suspended in the
center of the net mouth to measure volume of water filtered (see
Kramer et al., 1972, for further details).

The standard tow from 1951 through 1968 was an oblique haul
to 14 m depth (to 15 m of the bottom in shallow areas) designed
to filter a constant amount of water per depth interval (ca.
3m 3 /m of depth) over the vertical range of most ichthyoplankters.
Hauls were made at a ship speed of 1.5-2.0 knots and initiated by
clamping the net line to the towing cable with the 45 kg terminal

•^CalCOFI lines are arranged perpendicular to the coastline and
extend from the Canadian border (line 10) to below Cape San
Lucas, Baja California (line 157) . Stations were established
on the basis of a perpendicular to line 80 (off Pt. Conception)
at a point designated as station 60. Stations were plotted
seaward and shoreward from station 60 on each line. Cardinal
CalCOFI lines (those ending in "0") are 120 miles apart and
usually bracket two ordinal lines (ending in "3" or "7", so that
lines are 40 miles apart over most of the pattern. Cardinal
stations are 40 miles apart and typically these are separated by
a station number ending in "5" so that stations are 20 miles
apart out to station 90 on most lines. Stations are placed at
closer intervals near the coast and islands to accommodate these
features (see Kramer et al., 1972 for further details).

weight about 10-15 m below the surface. The net was lowered to
140 m depth by paying out 200 m of wire over a 4 minute period
(35 m of depth/min.). After fishing at depth for 3 seconds, the
net was retrieved at 20 m/min. (14 m depth/min. ) . The angle of
stray of the towing cable was recorded every 3 seconds and
maintained at 45° (±3°) by adjusting the ship speed and course.
After reaching the surface, the net was washed down and the
samples preserved in 5% formalin buffered with sodium borate.
Flowmeter readings were made at the beginning and end of each
tow. Detailed descriptions of gear and methods are given by
Ahlstrom (1953), Kramer et al. (1972), and Smith and Richardson
(1977) .

LABORATORY PROCEDURES

Laboratory processing began with the determination of a
displacement volume for each sample (methods described in Staff,
SPFI, 1953 and Kramer et al., 1972). Zooplankton volumes
(including ichthyoplankton) of samples collected in 1951 are
listed in Staff, SPFI (1952) and presented graphically in
Thrailkill (1956) and Smith (1971).

Sorting involved the removal of ichthyoplankton from the
sample and identification and separation of eggs and larvae of
selected species (see introduction) . Usually, each sample was
sorted completely; however, some of the samples were fractioned
into aliguots using a Folsom plankton splitter (McEwen et al.,
1954) prior to sorting. Several criteria were used to determine
whether a sample was fractioned: samples containing an abundance
of thaliacians and coelenterates and exceeding 150 ml in total
plankton volume were fractioned (to 50%, 25%, 12.5%, or 6.25%) to
approximate a reduced volume of 50 ml for sorting; samples with
an excessive guantity of fish eggs and/or larvae were
occasionally fractioned to expedite the sorting process in order
to meet scheduled deadlines. If the identified fraction of an
aliquot yielded rare or interesting species of fish larvae, the
remaining fraction was frequently sorted and identified with the
intent of finding additional specimens. Aliquot percentages for
fractioned samples from 1951 are listed in Table 1 under the
"Percent Sorted" column.

A "standard haul factor" (SHF) was calculated for each tow
to make them comparable and allow estimations of areal abundance.
This factor adjusts the number of eggs or larvae in a haul to the
number in 10 m of water strained per meter of depth fished. If
the vertical distribution of the species has been encompassed,
then the adjusted value is equivalent to the number under 10 m
of sea surface. The SHF is calculated for each haul by the
formula:

Personal communication, James R. Thrailkill, National Marine
Fisheries Service, Southwest Fisheries Center, La Jolla, CA.

SHF = 10 D
V

where D = depth of haul = cosine of the average angle
of stray of the towing cable multiplied by
cable length (m)

V = total volume of water (m ) strained

during the haul

V = R * a ' p

where R = total number of revolutions of the current
meter during the haul

a = area (m ) of the mouth of the net

p = length of column of water (m) needed to
produce one revolution of the current
meter.

Tow depth, volume of water strained, and standard haul factor are
listed in Table 1 for each tow taken during 1951. Detailed
descriptions of factors involved in calculating these values are
presented in Ahlstrom (1948), Kramer et al. (1972), and Smith and
Richardson (1977) .

IDENTIFICATION

Identification of ichthyoplankton species beyond those
separated during the sorting process was carried out by a
separate group of specialists. Ontogenetic stages of fishes are
inherently difficult to identify and this is further complicated
by the large number and diversity of species which contribute to
the ichthyoplankton of the California Current region. Most
identifications were accomplished by establishing ontogenetic
series on the basis of morphology, meristics, and pigmentation
and then identifying these series by relating them to known
metamorphic, juvenile, or adult stages with overlapping features
(Powles and Markle, 1984) . A total of about 118 taxa were
identified in 1951, with 63 taken to species, 26 to genus, 25 to
family, and 4 to order (after present corrections 132, 67, 29,
31, 5, respectively). Some of the developmental series
recognized at the time could not be assigned scientific names,
particularly in the Bathylagidae, Myctophidae, and
Pleuronectiformes. These were given descriptive names, which
later were changed to scientific names as they became known.
Knowledge of the ontogenetic stages and of the taxonomy of the
regional ichthyofauna have increased markedly over the years so
that we now identify twice the number of taxa known in 1951.

The task of producing a reliable and equitable
ichthyoplankton data base required extensive procedures to
verify, correct, and edit the original identifications. The

primary data source was the original identification sheets (see
Kramer et al., 1972, for examples); however, a critical resource
used in all phases of this process was the SWFC ichthyoplankton
collection in which the CalCOFI samples are archived. Throughout
the course of CalCOFI ichthyoplankton studies, samples have been
identified to the lowest taxon possible. In reviewing these
identifications for the data base, our approach has been
conservative and we have preserved those identifications and
counts which we could confirm, while correcting as many of the
errors as possible. During the coding of the identification
sheets, the "descriptive types" were assigned scientific names
and reexamined, if necessary. After computer entry, taxonomic
errors and inconsistencies in the data base were corrected and
the most obvious identification errors were corrected. Our
current knowledge of ichthyoplankton technigues coupled with a
precise understanding of the development of identification
competency in the program over the years allowed us to critically
judge the historical records. Identifications were changed to
different taxa, lumped to a higher taxonomic category, or given a
more precise taxonomic name. In many cases, identifications of a
taxon were inconsistent among cruises in a year, because of
varying competency of identifiers. These records were made
equitable by lumping to the higher taxonomic category to avoid
biases that could result in quantitative misinterpretations.

Next, statistical, seasonal, and geographic outliers were
identified, employing a series of graphic summaries and listings.
Examination of geographic outliers proved to be especially
effective because of our accumulated knowledge of species
distributions. In the course of examining samples for these
outliers, other identification errors were discovered and
eventually all taxa were scrutinized to some extent. Lastly,
certain taxa were reexamined in all samples for the entire
CalCOFI time series. These taxa were selected because of their
commercial, ecological, phylogenetic, or zoogeographic importance
or because taxonomic confusion was at the ordinal level. The
following is a list of the taxa for 1951 which received special
attention, with explanations and caveats intended to aid in
quantitative interpretations:

Anguilliformes - tentative and sporadic identifications to family
or lower taxon lumped to order.

Sardinops sagax - all specimens south of line 12 checked for
misidentif ication of Opisthonema spp.

Engraulidae - includes nearshore taxa (mostly Anchoa spp.) large
enough to separate from Engraulis mordax . Some nearshore
samples of small E. mordax may contain other anchovy genera,
but could not be differentiated.

Nansenia spp. - all specimens checked and identified as TV.

Candida or JV. crassa; all specimens of these species near
their range boundaries checked.

Bathylagus wesethi - all listings of B. nigrigenys were southern
specimens of B. wesethi.

Sternoptychidae - tentative and sporadic identifications of
hatchetf ishes to genus were lumped to family.

Bathophilus spp. - all specimens checked.

Tactostoma macropus - all specimens checked.

Scopelarchidae - tentative and sporadic identifications to genus
lumped to family.

Lampanyctus spp. - tentative and sporadic identifications to
species (mostly descriptive types) lumped to genus.

Diogenichthys atlanticus - all specimens at margins of range
checked.

Diogenichthys laternatus - all specimens at margins of range
checked.

Electrona rissoi - recognition of this species was inconsistent
and others may be included in Protomyctophum crockeri or
Myctophidae.

Hygophum spp. - all specimens reidentified to species; residuals
are small, poorly preserved specimens.

Myctophum aurolaternatum - all specimens of "Astronesthidae"
proved to be this species.

Protomyctophum crockeri - some samples on northern lines may
contain P . thompsoni , which was not identified at the
time.

Symbolophorus calif orniensis - all specimens south of line 120
checked for confusion with Hygophum spp., stemming from
descriptive names.

Bregmaceros spp. - all gadiform types (see Index) , except
Merluccius productus and Macrouridae, reexamined.

Ophidiiformes - this category did not exist originally and

ophidiiform larvae were included in Brosmophysis marginata ,
Carapidae, "Otophidium" , "Zoarcidae", and "blenny" ; identi-
fications of B. marginata and Carapidae proved to be mostly
correct and "Zoarcidae" to be a yet unidentified ophidiiform
species; all "Otophidium" and "blenny" were reexamined and
the former included Ophidion scrippsae , Chilara taylori and
other ophidiiform taxa (moved to order) ; "blenny" contained
0. scrippsae, C. taylori, and other ophidiiform taxa in
addition to true blennioids.

Ceratioidei - identifications of this group were inconsistent and
additional specimens may be in the unidentified fish larva
category.

Hemiramphidae - all specimens checked.

Trachipteridae - tentative and sporadic identifications to genus
were lumped to family.

Melamphaes spp. - all identifications ascribed to Melamphaidae
were reexamined and assigned to genus (Melamphaes ,
Poromitra) or species (Scopelogadus bispinosus) .

Cottidae - some samples may include specimens of Scorpaenichthys
marmoratus , hexagrammids (e.g., Oxylebius pictus , Zaniolepis
spp.), and some blennioids (e.g., Hypsoblennius spp.).

Hexagrammidae - primarily Hexagrammos spp.

Scorpaenidae - now includes genera (Pontinus , Scorpaena ,

Scorpaenodes) other than Sebastes ; some of these other
genera remain in Sebastes , particularly on southern lines.

Sebastes spp. - in addition to other scorpaenid genera,
category includes some misidentif ied Sebastolobus spp.,
Prionotus spp., serranids, scombrids, and other spiny-
headed shorefishes, particularly in samples south of
line 120.

Blennioidei - this is the residual of the completely reexamined
"blenny" category, which also contained various misidenti-
fied ophidiiforms, and is now restricted to members of
northern stichaeioid families and true blennioids (other
than Hypsoblennius spp.) in the southern part of the
pattern) .

Hypsoblennius spp. - some specimens remain in Cottidae.

Clinidae - some specimens remain in Cottidae or unidentified fish
larva category.

Labridae - tentative and sporadic identifications to genus were
lumped to family.

Chromis punctipinnis - records south of about line 12 may
include other pomacentrid taxa.

Mugil spp. - all specimens checked.

Apogonidae - all specimens checked.

Carangidae - all specimens checked; tentative and sporadic
identifications to genus or species (except Trachurus
symmetricus) were lumped to family.

Gerreidae - larvae of this family and other shorefishes (e.g.,
Haemulidae, Girella nigricans , Caulolatilus princeps ,
Mullidae, Priacanthidae) were not identified and may be in
the unidentified fish larva category or may be misidentif ied
as Sebastes spp. , Cottidae, etc.

Sciaenidae - this family is underrepresented and some specimens
may be in the unidentified fish larva category or may have
been misidentif ied as Sebastes spp. , Serranidae, etc.

Scombridae - all larvae identified to this family or constituent
taxa (except Scomber japonicus) were reexamined and reas-
signed; underrepresentation or absence of these taxa may be
attributed to misidentif ication or they may be in the uni-
dentified fish larva category.

Nomeidae - absence of this family attributed to misidentif ication
or placement in unidentified fish larva category.

Pleuronectiformes - all available specimens of this category

(originally called "flatfish") were examined and reidenti-
fied; residuals are small, poorly preserved specimens.

Bothidae - all specimens examined and reassigned; most were

assigned to various paralichthyid genera or to Bothus spp.

Citharichthys spp. - tentative and sporadic identifications to

species were lumped to genus, which also includes Etropus
spp. and some other flatfish taxa (e.g., Parophrys vetulus) .

Hippoglossina spp. - all specimens of this genus (originally

called "pigmented bothid") were examined and those not
assigned to H. stomata were left at the genus (probably are
H. tetropthalmus) .

Paralichthys spp. - all specimens of this genus were examined and
most were assigned to P. calif ornicus or Xystreurys
liol epi s .

Syacium ovale - all specimens examined (originally called "spiny-
headed bothid") .

Xystreurys liolepis - originally misidentif ied as Paralichthys
calif ornicus; all specimens reidentif ied.

Glyptocephalus zachirus - all specimens examined.

Microstomus pacificus - all specimens examined.

Pleuronichthys spp. - all larvae of this genus and constituent

species were examined and assigned to species; residuals are
small, poorly preserved specimens.

Psettichthys melanost ictus - absence of this species may be

explained by misidentif ication with other flatfish species

(e.g., Lyopsetta exilis) which we did not reexamine
systematically.

COMPUTER ENTRY AND EDITING

Each taxon on the original identification sheets was given a
3-digit code based on the list of codes in Haight et al. (1979).
Taxon codes and counts from these sheets were keypunched by
cruise and station, along with pertinent station and tow data and
entered into the VAX 11/780 computer at the University of
California, San Diego Computing Center. After entries were
completed for an entire year, print-out listings of taxa and
counts on each station were compared with the original data
sheets to eliminate keypunch errors. Next, data in the file were
cross-checked with data on an existing file which contained:
station and tow data; numbers of eggs of sardine, anchovy, and
saury (Cololabis saira) ; numbers of larvae of sardine, anchovy,
hake, jack mackerel, and Pacific mackerel; total number of fish
eggs; and total number of fish larvae.

Discrepancies in ichthyoplankton data in these two files
were corrected by inspecting original records from the sorting
laboratory, the original ichthyoplankton identification sheets,
and the samples themselves. Station and tow data discrepancies
between the two files were corrected by reviewing ships logs and
deck tow sheets, original records from the sorting laboratory,
cruise announcements, publications, header information on the
ichthyoplankton identification sheets, and station plots
generated for each cruise. Eventually all station and tow data
were checked by comparing these sources.

The corrected ichthyoplankton data base was then examined
statistically and outliers were found and checked as above.
Distributional plots were then prepared for each taxon and these
were checked by reviewing the data sources mentioned above and by
examining archived specimens. A listing of each taxon by station
(Table 4) was produced, which became the primary document for
subseguent checks. Misidentif ications found in geographic
outlier checks and other misidentif ications and data problems
discovered in the course of examining archived samples resulted
in several iterations of Table 4. Finally, totals in Table 4
were checked against annual summaries of incidence and abundance
(Tables 2 and 3). Ecological analyses of the data (Moser et al.,
1987) were conducted concurrently with editing procedures and
provided cross-checks that allowed correction of errors.

SPECIES SUMMARY

Larvae of Pacific hake (Merluccius productus) represented
24% of all fish larvae taken on CalCOFI cruises during 1951 and
numbered twice as many as northern anchovy (Engraulis mordax) ,
the next most abundant species (Tables 2, 3). Hake incidence was
comparatively low (11th) as was anchovy incidence (7th) ,

10

indicating relatively large sample sizes. The next most abundant
species, Vinciguerria lucetia , a midwater gonostomatid, was
widespread in samples from the southern part of the CalCOFI
pattern and ranked 4th in occurrence. Jack mackerel larvae
ranked 4th in abundance and 8th in occurrence. Larvae of
Sebastes spp. , a composite of about 70 species, ranked 5th in
abundance and 1st in occurrence. The next most abundant were
three midwater species, a deepsea smelt (Leuroglossus stilbius)
and two lanternf ishes (Stenobrachius leucopsarus and Triphoturus
mexicanus) , ranking 6th, 7th, and 8th, respectively. The latter
ranked 2nd in occurrence. Sardine larvae ranked 9th in
abundance but only 21st in occurrence, indicating large sample
sizes. The 10th ranked taxon was the sanddab genus
Citharichthys , a composite of about 6 species whose distributions
in aggregate cover most of the CalCOFI pattern. These 10 top-
ranking taxa contributed 84.2% of all larvae taken during 1951.
The remaining 15.8% is represented by 124 taxa. Of the 10 taxa,
4 were midwater species, 3 were coastal demersal species or
generic groupings, and 3 were coastal pelagic species.

EXPLANATION OF TABLES

Table 1 - This table lists the pertinent station and tow data for
1951, the volume of water filtered and standard haul
factor for each tow, the percent of sample sorted, and
the total numbers of fish eggs and larvae. CalCOFI
cruises are designated by four digits; the first two
indicate the year and the second two the month.
Within each cruise the data are listed in order of
increasing line and station number (southerly and
seaward directions) ; the order of station occupancy is
shown on the station charts (Figures 2-13). Stations
are designated by two groups of digits; the first set
indicates the line and decimal fraction, if necessary,
and the second set indicates the station on the line.
Decimal fractions were not used in 1951. Time is
listed as Pacific Standard Time at the start of each
tow in 24-hour designation. Methods for determining
tow depth, volume of water strained, standard haul
factor, and percent sorted were described in the
methods section. The values for total fish eggs and
larvae represent raw counts (unadjusted for percent
sorted or standard haul factor) . Ship codes are as
follows: BD, Black Douglas; CR, Crest; HO, Horizon;
PT, Paolina T; YE, Yellowfin; 47, N. B. Scofield .

Table 2 - This table lists pooled occurrences of all larval fish
taxa taken during 1951 in ranked order.

Table 3 - This table lists pooled counts of all larval fish taxa
taken during 1951 in ranked order. Numbers are
adjusted for percent sorted and standard haul factors.

11

Table 4 - This table gives numbers of fish larvae
listed by station and month. Counts a
percent of sample sorted and standard
Average values are given for stations
than once during a month. See Table 1
tow data and Table 6 for listing of
double occupancies during a month,
listed in "phylogenetic" sequence modif
(1984) . Subtaxa within each order
alphabetically. Page numbers for each
in the index at the end of the report.

for each taxon,

re adjusted for

haul factor.

occupied more
for station and

stations with

The orders are

ied from Nelson

are listed

taxon are given

Table 5 - This table is a summary of pooled occurrences of all
larval fish taxa taken on CalCOFI surveys from 1951 to
1960. Taxa are listed in the same order as in Table 4.

Table 6 - List of stations which were occupied twice in one month
during 1951. Ship codes are given in explanation for
Table 1.

ACKNOWLEDGMENTS

Elbert Ahlstrom, David Kramer, Robert Counts, Orville Ball,
and Robert Wisner originally identified larvae from CalCOFI
cruises of 1951. Ronald Whyte coded each larval fish taxon or
type and Rita Ford entered them into the computer. Debby Snow
efficiently assisted in all aspects of data editing and
retrieval. Cindy Meyer, Larry Zins, and James Ryan provided
programming assistance. Dorothy Roll designed the CalCOFI data
acquisition system and provided data processing support. Ken
Raymond and Roy Allen helped with graphics and production of the
report. Lorraine Prescott and Diane Forsythe prepared the
manuscript for printing. Paul Smith determined statistical
outliers, provided assistance during geographical outlier checks
and offered helpful suggestions throughout the project. Izadore
Barrett, Director of the Southwest Fisheries Center and Reuben
Lasker, Chief, Coastal Fisheries Resources Division, SWFC,
provided the support critical to the completion of the project.
James Thrailkill planned CalCOFI surveys and supervised cruises,
data handling, and plankton sorting from 1949 to 1986 and is
largely responsible for the high quality of these operations.
Without the vision and direction of Elbert Ahlstrom and Elton
Sette and the dedicated efforts of the many people who collected,
processed, and analyzed the samples, this data base would not
exist.

12

LITERATURE CITED

Ahlstrom, E. H. 1948. A record of pilchard eggs and larvae
collected during surveys made in 1939 to 1941. U.S. Fish
Wildl. Serv. SSRF 54, 82 p.

Ahlstrom, E. H. 1953. Pilchard eggs and larvae and other fish
larvae, Pacific Coast - 1951. U.S. Fish Wildl. Serv. SSRF
102, 55 p.

Ahlstrom, E. H. 1969. Distributional atlas of fish larvae in the
California Current region: jack mackerel, Trachurus
symmetricus , and Pacific hake, Merluccius productus , 1951
through 1966. CalCOFI Atlas No. ll:xi + 187 p.

Ahlstrom, E. H. , H. G. Moser, and E. M. Sandknop. 1978.
Distributional atlas of fish larvae in the California
Current region: Rockfishes, Sebastes spp., 1950 through
1975. CalCOFI Atlas No. 26: xxi + 178 p.

Haight, C. A., H. G. Moser, and P. E. Smith. 1979. Data entry
programs: CalCOFI. II. Fish eggs and larvae identification
sheet. National Marine Fisheries Service, Southwest
Fisheries Center, La Jolla, Admin. Rept. No. LJ-79-25.

Kramer, D. 1970. Distributional atlas of fish eggs and
larvae in the California current region: Pacific sardine,
Sardinops caerulea (Girard) , 1951 through 1966. CalCOFI
Atlas No. 12:vi + 277 p.

Kramer, D. and E. H. Ahlstrom. 1968. Distributional atlas of
fish larvae in the California Current region: Northern
anchovy, Engraulis mordax (Girard) , 1951 through 1965.
CalCOFI Atlas No. 9: xi + 269 p.

Kramer, D. , M. Kalin, E. G. Stevens, J. R. Thrailkill, and J. R.
Zweifel. 1972. Collecting and processing data on fish eggs
and larvae in the California Current Region. NOAA Tech.
Rep. NMFS Circ. 370, 38 p.

McEwen, G. F. , M. W. Johnson, and T. R. Folsom. 1954. A
statistical analysis of the performance of the Folsom
Plankton Sample Splitter, based on test observations. Arch.
Meteor. Geophys. Bioklim. Ser. A, 7:502-527.

Moser, H. G. , P. E. Smith, and L. E. Eber. 1987. Larval fish
assemblages in the California Current region during 1954-
1960, a period of dynamic environmental change. CalCOFI
Rep. 28.

Nelson, J. S. 1984. Fishes of the world. John Wiley and Sons,
N.Y. , 523 p.

13

Powles, H. and D. F. Markle. 1984. Identification of larvae, p.
31-3 3. In: Ontogeny and systematics of fishes. H. G.
Moser, W. J. Richards, D. M. Cohen, M. P. Fahay, A. W.
Kendall, Jr., and S. L. Richardson (eds.). Spec. Publ.
No. 1. Amer. Soc. Ichthyol. Herpetol .

Reid, J. L. , Jr., R. S. Arthur, and E. B. Bennett, (eds.).
1963. Oceanic observations of the Pacific: 1951. Univ.
Calif. Press, Berkeley, 598 p.

Smith, P. 1971. Distributional atlas of zooplankton volume in
the California Current region, 1951 through 1966. CalCOFI
Atlas No. 13: xvi + 144 p.

Smith, P. E. and S. L. Richardson. 1977. Standard techniques
for pelagic fish egg and larva surveys. FAO Fish. Tech.
Pap. No. 175, 100 p.

Staff, South Pacific Fishery Investigations. 1952. Zooplankton
volumes off the Pacific Coast, 1951. U.S. Fish Wildl. Serv.
SSRF 73, 37 p.

Staff, South Pacific Fishery Investigations. 1953. Zooplankton
volumes off the Pacific Coast, 1952. U.S. Fish Wildl. Serv.
SSRF 100, 41 p.

Thrailkill, J. R. 1956. Relative areal zooplankton abundance
off the Pacific coast. U.S. Fish Wildl. Serv. SSRF 188,
85 p.

14

Figure 1. Composite arrangement of diagrammatic charts showing
areas sampled on each CalCOFI cruise during 1951.

15

r

40°

30°

20°

J I L

J L.

J L

J I L

Figure 2. Station pattern for CalCOFI Cruise 5101 showing
tracks for each vessel. Stations with plankton tows
only are indicated by a dot; those with plankton tows
and hydrographic measurements are shown by a dot and
circle. Modified from charts in Oceanic Observations
of the Pacific (Reid et al . , 1963) to include only
those stations listed in Table 1 of this report.

16

125°
-1 1 1 1—5

120° 115° 110°
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120°

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Figure 3. Station pattern for CalCOFI Cruise 5102. Symbols as
in Figure 2 .

17

40'

35"

30°

25"

20°

i 1 1 r

CRUISE 5103

BLACK DOUGLAS

J L

i i i

J L

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— 20°

125°

120°

115°

110°

Figure 4. Station pattern for CalCOFI Cruise 5103
in Figure 2 .

Symbols as

18

40'

35 c

30°

25'

20'

HORIZON

J I L

' ' ' L

J L

i i i

20°

125"

120°

no-

Figure 5. Station pattern for CalCOFI Cruise 5104. Symbols as
in Figure 2 .

19

40'

35'

30°

25 c

20'

i 1 r

CRUISE 5105

HORIZON

35°

J L

J I L

J I L

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125°

120°

110°

Figure 6. Station pattern for CalCOFI Cruise 5105. Symbols as
in Figure 2 .

20

i 1 r

CRUISE 5106

40°

35°

30°

25°

20°

J L

I I I I

I I I

20°

125°

120°

115°

110°

Figure 7. Station pattern for CalCOFI Cruise 5106,
in Figure 2 .

Symbols as

21

125°

Figure 8. Station pattern for CalCOFI Cruise 5107. Symbols as
in Figure 2.

22

40°

35°

30°

25°

PAOLINA-T

20°

J L

i i i i

— 20°

125°

120°

115°

Figure 9. Station pattern for CalCOFI Cruise 5108
in Figure 2.

Symbols as

23

Figure 10. Station pattern for CalCOFI Cruise 5109
in Figure 2 .

Symbols as

24

40'

35'

30°

25'

20'

J ' '

125°

J L

120°

J L

20°

Figure 11. Station pattern for CalCOFI Cruise 5110. Symbols as
in Figure 2 .

25

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120° 115° 110°
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Figure 12. Station pattern for CalCOFI Cruise 5111. Symbols as
in Figure 2.

26

40° —

t r

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30°

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20°

CRUISE 5112

YELLOWHN

HORIZON

J L

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Figure 13. Station pattern for CalCOFI Cruise 5112. Symbols as
in Figure 2 .

27

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64

TABLE 2. Pooled occurrences of fish larvae taken during CalCOFI
cruises in 1951.

Rank Taxon Occurrences

1 Sebastes spp. 600

2 Triphoturus mexicanus 589

3 Lampanyctus spp. 57 6

4 Vinciguerria lucetia 532

5 Citharichthys spp. 428

6 Leuroglossus stilbius 402

7 Engraulis mordax 394

8 Trachurus symmetricus 37 2

9 Protomyctophum crockeri 370

10 Stenobrachius leucopsarus 369

11 Merluccius productus 3 51

12 Tarletonbeania crenularis 306

13 Bathylagus wesethi 259

14 Cyclothone spp. 253

15 Diogenichthys laternatus 2 30

16 Disintegrated fish larva 229

17 Melamphaes spp. 221

18 Symbolophorus calif orniensis 206

19 Unidentified fish larva 187

20 Paralepididae 169

21 Sardinops sag ax 167

22 Bathylagus ochotensis 153

23 Ceratoscopelus townsendi 140

24 Icichthys lockingtoni 125

25 Gobiidae 116
2 5 Diaphus spp. 116

27 Diogenichthys atlanticus 109

28 Myctophidae 99

29 Stomias atriventer 96

30 Labridae 74

31 Ophidiif ormes 68

32 Scomber japonicus 59
32 Scopelarchidae 59
34 Argentina sialis 55
34 Chauliodus macouni 55

36 Cololabis saira 53

37 Lyopsetta exilis 51

38 Nansenia crassa 50

39 Gonichthys tenuiculus 49

40 Idiacanthus antrostomus 48

41 Hygophum atratum 47

42 Symphurus spp. 4 5

43 Synodus spp. 41

44 Lampadena urophaos 3 9

45 Sternoptychidae 38

46 Chromis punctipinnis 37

47 Anguilliformes 35

65

TABLE 2. (Cont . )

Rank Taxon Occurrences

48 Trachipteridae 32

49 Myctophum nitidulwn 30

50 Nansenia Candida 29
50 Bygophum spp. 29
50 Tetragonurus cuvieri 29
5 3 Microstomus pacificus 28

54 Etrumeus acuminatus 2 5

55 Cottidae 24
55 Sebastolobus spp. 24
55 Prionotus spp. 24
55 Chiasmodontidae 24

59 Trichiuridae 23

60 Microstoma microstoma 21

61 Serranidae 20
61 Tactostoma macropus 20

63 Loweina rara 19

64 Hypsoblennius spp. 18
64 Paralichthys calif ornicus 18
66 Hygophum reinhardtii 17
66 Ophidion scrippsae 17
66 Pleuronichthys coenosus 17
69 Ichthyococcus spp. 16
69 Notoscopelus resplendens 16
69 Aristostomias scintillans 16
7 2 Electrona rissoi 15
72 Carangidae 15
74 Peprilus simillimus 14
74 Sphyraena argentea 14
7 4 Pleuronichthys spp. 14

77 Hippoglossina stomata 13

78 Sciaenidae 12
78 Bathylagus pacificus 12
78 Glyptocephalus zachirus 12
81 Scorpaenidae 10
81 Diogenichthys spp. 10
83 Auxis spp. 9
83 Brosmophycis marginata 9
83 Physiculus spp. 9
83 Pleuronectif ormes 9
83 Medialuna californiensis 9
88 Diplophos taenia 8
88 Exocoetidae 8

90 Clinidae 7

91 Scorpaenichthys marmoratus 6
91 Myctophum aurolaternatum 6
91 Chilara taylori 6
94 Notolychnus valdiviae 5
9 4 Syngnathus spp. 5

66

TABLE 2. (Cont. )

Rank Taxon Occurrences

94 Syacium ovale 5

94 Hemiramphidae 5

94 Macrouridae 5

99 Scopelogadus bispinosus 4

99 Bathophilus spp. 4

99 Cyclopteridae 4

99 Brama spp. 4

99 Pleuronichthys decurrens 4

104 Albula wipes 3

104 Xystreurys liolepis 3

104 Pleuronichthys verticalis 3

104 Ceratioidei 3

104 Bothus spp. 3

109 Tetraodontidae 2

109 Mugil spp. 2

109 Alepocephalidae 2

109 Carapidae 2

109 Gempylidae 2

109 Atherinidae 2

109 Porichthys spp. 2

109 Agonidae 2

109 Bregmaceros spp. 2

109 Blennioidei 2

119 Hippoglossina spp. 1

119 Anotopterus pharao 1

119 Hexagrammidae 1

119 Poromitra spp. 1

119 Opisthonema spp. 1

119 Apogonidae 1

119 Macroramphosus gracilis 1

119 Bathylagus milleri 1

119 Uranoscopidae 1

119 Antennariidae 1

119 Moridae 1

119 Icosteus aenigmaticus 1

119 Scomberomorus spp. 1

119 Balistidae 1

119 Aulopus spp. 1

119 Pleuronichthys ritteri 1

67

TABLE 3. Pooled numbers of fish larvae taken during CalCOFI

cruises in 1951. Counts are adjusted for percent of
sample sorted and standard haul factor (see text).

Rank Taxon Count

1 Merluccius productus 6 2 594

2 Engraulis mordax 29 372

3 Vinciguerria lucetia 2 3308

4 Trachurus symmetricus 1917 5

5 Sebastes spp. 184 9 5

6 Leuroglossus stilbius 17696

7 Stenobrachius leucopsarus 16 248

8 Triphoturus mexicanus 13067

9 Sardinops sag ax 110 58

10 Citharichthys spp. 8411

11 Diogenichthys laternatus 3803

12 Lampanyctus spp. 3516

13 Tarletonbeania crenularis 3180

14 Bathylagus wesethi 2578

15 Unidentified fish larva 2196

16 Diaphus spp. 2 48

17 Disintegrated fish larva 1778

18 Cyclothone spp. 1547

19 Protomyctophum crockeri 1524

20 Synodus spp. 1341

21 Symbolophorus californiensis 116 8

22 Ceratoscopelus townsendi 1083

23 Bathylagus ochotensis 1070

24 Scomber japonicus 10 20

25 Ophidiiformes 1012

26 Symphurus spp. 908

27 Jcichthys lockingtoni 87 9

28 Prionotus spp. 867

29 Melamphaes spp. 723

30 Paralepididae 592

31 Diogenichthys atlanticus 585

32 Gobiidae 465

33 Etrumeus acuminatus 423

34 Labridae 391

35 Myctophidae 350

36 Stomias atriventer 34 2

37 Carangidae 322

38 Argentina sialis 310

39 Chromis punctipinnis 299

40 Serranidae 197

41 Opisthonema spp. 19 5

42 Scopelarchidae 188

43 Ophidion scrippsae 176

44 Lyopsetta exilis 172

45 Anguilliformes 169

68

TABLE 3. (Cont. )

Rank Taxon Count

46 Idiacanthus antrostomus 158

47 Hygophum atratum 154

48 Cololabis saira 151

49 Lampadena urophaos 147

50 Nansenia crassa 146

51 Gonichthys tenuiculus 143

52 Chauliodus macouni 137

53 Sphyraena argentea 119

54 Tetragonurus cuvieri 117

55 Cottidae 104

56 Microstomus pacificus 102

57 Sternoptychidae 101

58 Tactostoma macropus 97

59 Trachipteridae 95

60 Paralichthys californicus 94

61 Auxis spp. 93

62 Bathylagus pacificus 92

63 Notoscopelus resplendens 91

64 Nansenia Candida 90

65 Hygophum spp. 84

66 Trichiuridae 79

67 Physiculus spp. 77

68 Sebastolobus spp. 7 2

69 Microstoma microstoma 69

70 Myctophum nitidulum 67

71 Sciaenidae 66

72 Chiasmodontidae 61
7 3 Peprilus simillimus 59
74 Diogenichthys spp. 57

7 5 Glyptocephalus zachirus 54

76 Pleuronichthys spp. 52

77 Exocoetidae 50

78 Hypsoblennius spp. 47

79 Hygophum reinhardtii 44
79 Electrona rissoi 44

81 Loweina rara 43

82 Hippoglossina stomata 38

8 2 Ichthyococcus spp. 38
84 Pleuronichthys coenosus 36
84 Scorpaenidae 36

86 Aristostomias scintillans 31

87 Moridae 29

88 Diplophos taenia 28
88 Myctophum aurolaternatum 28

90 Pleuronectif ormes 26

91 Brosmophycis marginata 23
91 Clinidae 23
93 Hemiramphidae 20

69

TABLE 3. (Cont. )

Rank Taxon Count

94 Medialuna calif orniensis 19

9 5 Syngnathus spp. 18

96 Albula vulpes 17

97 Syacium ovale 15
97 Brama spp. 15
99 Chilara taylori 14
99 Scorpaenichthys marmoratus 14
101 Bathophilus spp. 12
101 Notolychnus valdiviae 12

103 Macrouridae 11

104 Xystreurys liolepis 10

10 5 Scopelogadus bispinosus 9
106 Bothus spp. 8
106 Cyclopteridae 8
106 Pleuronichthys decurrens 8
109 Ceratioidei 7
109 Icosteus aenigmaticus 1
111 Carapidae 6
111 Tetraodontidae 6
111 Pleuronichthys verticalis 6
114 Atherinidae 5
114 Porichthys spp. 5
116 Mugil spp. 4
116 Bregmaceros spp. 4
116 Blennioidei 4
116 Agonidae 4
116 Gempylidae 4
121 Scomberomorus spp. 3
121 Antennariidae 3
121 Poromitra spp. 3
121 Uranoscopidae 3
121 Alepocephalidae 3
121 Bathylagus milleri 3
121 Macroramphosus gracilis 3
128 Hexagrammidae 2
128 Apogonidae 2
128 Anotopterus pharao 2
128 Pleuronichthys ritteri 2
128 Balistidae 2
128 Hippoglossina spp. 2
128 Aulopus spp. 2

Total 260740

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190

TABLE 6. List of stations which were occupied twice in
one month during 1951.

Station

Mon

110.0

35.0

1

83.0

55.0

5

83.0

70.0

5

87.0

35.0

5

87.0

40.0

5

93.0

30.0

5

123.0

40.0

5

123.0

50.0

5

123.0

60.0

5

127.0

50.0

5

127.0

60.0

5

133.0

30.0

5

133.0

40.0

5

133.0

50.0

5

137.0

35.0

5

130.0

35.0

6

130.0

40.0

6

130.0

50.0

6

130.0

60.0

6

137.0

50.0

6

70.0

60.0

8

70.0

70.0

8

70.0

80.0

8

73.0

60.0

8

77.0

50.0

8

77.0

55.0

8

77.0

65.0

8

120.0

25.0

8

120.0

30.0

8

120.0

35.0

8

120.0

45.0

8

120.0

60.0

8

120.0

70.0

8

80.0

51.0

9

80.0

55.0

9

80.0

60.0

9

80.0

70.0

9

80.0

80.0

9

80.0

90.0

9

80.0

100.0

9

83.0

55.0

9

85.0

38.0

9

85.0

40.0

9

85.0

50.0

9

90.0

70.0

9

123.0

37.0

9

123.0

40.0

9

Stat

ion

Mont

93.0

27.0

10

93.0

30.0

10

93.0

50.0

10

97.0

30.0

10

97.0

32.0

10

97.0

40.0

10

97.0

50.0

10

100.0

29.0

10

100.0

30.0

10

100.0

40.0

10

103.0

30.0

10

103.0

35.0

10

103.0

40.0

10

107.0

32.0

10

107.0

35.0

10

107.0

40.0

10

63.0

52.0

11

63.0

55.0

11

67.0

50.0

11

67.0

55.0

11

67.0

65.0

11

70.0

51.0

11

70.0

70.0

11

70.0

80.0

11

73.0

50.0

11

73.0

60.0

11

77.0

50.0

11

120.0

60.0

11

120.0

70.0

11

191

INDEX

This index lists taxa included in Table 4 with their page
numbers .

Page

Elopiformes

Albulidae

Albula vulpes 71

Anguilliformes 71

Clupei formes

Clupeidae

Etrumeus acuminatus 72

Opi sthonema spp

Sardinops sagax 72

Engraulidae

Engraulis mordax 74

Salmoni formes

Alepocephalidae 77

Argentinidae

Argentina sialis

Microstoma microstoma 78

Nansenia Candida 78

Nansenia crassa 79

Bathylagidae

Bathylagus milleri 80

Bathylagus ochotensis 80

Bathylagus pacificus 82

Bathylagus wesethi 82

Leuroglossus stilbius 85

Stomiif ormes

Gonostomatidae

Cycl othone spp 87

Diplophos taenia 90

Ichthyococcus spp 90

Vinciguerria lucetia 91

Sternoptychidae 94

Stomiatoidea

Chauliodontidae

Chauliodus macouni 95

Idiacanthidae

Idi acanthus antrostomus 96

Malacosteidae

Ari stostomi as scintillans 97

Melanostomiidae

Bathophilus spp 97

Tactostoma macropus 97

Stomiidae

Stomias atriventer 98

192

Page
Myctophi formes
Alepisauroidei

Anotopteridae

Anotopterus pharao 99

Paralepididae 99

Aulopoidei

Aulopidae

Aulopus spp 102

Chloropthalmoidei

Scopelarchidae 102

Myctophoidei

Myctophidae 103

Lampanyctinae

Ceratoscopel us townsendi 104

Diaphus spp 106

Lampadena urophaos 108

Lampanyctus spp 109

Notolychnus valdiviae 113

Notoscopelus resplendens 114

Stenobrachius leucopsarus 114

Triphoturus mexicanus 117

Myctophinae

Di ogeni chthys spp 121

Diogenichthys atlanticus 121

Diogenichthys laternatus 12 2

Electrona rissoi 124

Gonichthys tenuiculus 12 5

Hygophum spp 126

Hygophum atratum 12 6

Hygophum reinhardtii 127

Loweina rara 12 7

Myctophum aurolaternatum 128

Myctophum nitidulum 128

Protomyctophum crockeri 129

Symbolophorus calif orniensis 132

Tarletonbeania crenularis 134

Synodontoidei

Synodontidae

Synodus spp 137

Gadiformes

Bregmacerotidae

Bregmaceros spp 137

Merlucciidae

Merl ucci us productus 137

Moridae 141

Physi cul us spp 141

Macrouridae 141

Ophidii formes 141

Bythitidae

Brosmophysis marginata 142

Carapidae 142

193

Page
Ophidiidae

Chilara taylori 14 3

Ophidion scrippsae 143

Bat racoidi formes

Batracoididae

Pori ch thys spp 143

Lophii formes
Antennarioidei

Antennariidae 143

Ceratioidei 143

Beloniformes

Exocoetidae 144

Hemiramphidae 144

Scomberesocidae

Cololabis saira 144

Atherini formes

Atherinidae 145

Lampri formes

Trachipteridae 145

Beryci formes

Melamphaidae

Melamphaes spp 14 6

Poromi tra spp 149

Scopelogadus bispinosus 149

Syngnathi formes

Macroramphosidae

Macroramphosus gracilis 149

Syngnathidae

Syngnathus spp 149

Scorpaeni formes
Cottoidei

Agonidae 149

Cottidae 149

Scorpaeni chthys marmoratus 150

Cyclopteridae 150

Hexagrammidae 150

Scorpaenoidei

Scorpaenidae 150

Sebastes spp 151

Sebastolobus spp 155

Triglidae

Prionotus spp 155

Perciformes

Blennioidei 156

Blenniidae

Hypsoblennius spp 156

Clinidae 156

Gobioidei

Gobiidae 157

194

Page
Icosteoidei

Icosteidae

Icosteus aenigmaticus 158

Labroidei

Labridae 159

Pomacentridae

Chromis punctipinnis 160

Mugiloidei

Mugilidae

Mugil spp 161

Percoidei

Apogonidae 161

Bramidae

Brama spp 161

Carangidae 161

Trachurus symmetri cus 161

Kyphosidae

Medialuna calif orniensis 164

Sciaenidae 165

Serranidae 165

Scombroidei

Gempylidae 166

Scombridae

Auxis spp 166

Scomber japonicus 166

Scomberomorus spp 167

Trichiuridae 167

Sphyraenoidei

Sphyraenidae

Sphyraena argentea 168

Stroma teoidei

Centrolophidae

Icichthys lockingtoni 168

Stromateidae

Peprilus simillimus 170

Tetragonuridae

Tetragonurus cuvieri 170

Trachinoidei

Chiasmodontidae 171

Uranoscopidae 171

Pleuronectiformes 171

Pleuronectoidei
Bothidae

Bothus spp 171

Paralichthyidae

Citharichthys spp 172

Hippoglossina spp 175

Hippoglossina stomata 17 5

Paralichthys calif ornicus 175

Syacium ovale 17 6

Xystreurys liolepis 17 6

195

Page
Pleuronectidae

Glyptocephalus zachirus 17 6

Lyopsetta exilis 176

Microstomias pacificus 177

Pleuronichthys spp 178

PI euroni chthys coenosus 178

PI euroni chthys decurrens 179

Pi euroni chthys ritteri 1 79

Pleuronichthys verticalis 179

Soleoidei

Cynoglossidae

Symphurus spp 179

Tetraodonti formes

Balistidae 180

Tetraodontidae 180

Disintegrated fish larva 180

Unidentified fish larva 183

196

RECENT TECHNICAL MEMORANDUMS

Copies of this and other NOAA Technical Memorandums are available from the National Technical Informa-
tion Service, 5285 Port Royal Road, Springfield, VA 22167. Paper copies vary in price. Microfiche copies cost
$4.50. Recent issues of NOAA Technical Memorandums from the NMFS Southwest Fisheries Center are listed
below:

NOAA TM-NMFS SWFC

69 Effects of Tropical Tuna Fisheries on non-target species.
G.T. SAKAGAWA

(February 1 987)

70 The Hawaiian monk seal on Laysan Island: 1984.
T.C. JOHANOS, A.K.H. KAM, and R.G. FORSYTH
(March 1987)

71 Preliminary assessment of habitat utilization by Hawaiian green
turtles in their resident foraging pastures.

G.H. BALAZS, R.G. FORSYTH, and A.K.H. KAM
(March 1987)

72 Forces of change in Hawaii's aku (skipjack tuna) industry, 1986-
workshop summary.

C.H. BOGGS and S.G. POOLEY
(April 1987)

73 United States North Pacific Albacore Fishery 1 961 -1 980.
A.P. MAJORS

(April 1987)

74 Abundance of zooplankton species in California coastal waters
during April 1981, February 1982 and March 1985.

A. ALVARINO and C.A. KIMBRELL
(June 1987)

75 Data report on the vertical distribution of the eggs and larvae of
northern anchovy, Engraulis mordax, at two stations in the Southern
California Bight, March-April 1980.

T. POMMERANZ and H.G. MOSER
(July 1987)

76 Report of a marine mammal survey of the Eastern Tropical Pacific
aboard the research vessel David Starr Jordan, July 29-December 5,
1986.

R.S. HOLT and S.N. SEXTON
(August 1987)

77 Report of a marine mammal survey of the Eastern Tropical Pacific
aboard the research vessel McArthur, July 29-December 6, 1986.
R.S. HOLT and A. JACKSON

(August 1987)

78 Results of the Bay Area Sportfish Economic Study (BASES)
C.J. THOMSON and D.D. HUPPERT

(August 1987)